Grain size characteristics of the sediment and the flow stage characteristics of the river are the two most important factors influencing sedimentation in the channel zone. Climate is important in its influence on floodplain (overbank) sediment.

Supra-bar platform (upper) parts of mature bars from a wide range of climatic conditions consistently show a sequence in the development of contained sedimentary structures which is related to the flow stage. At high stage, in rivers where the bed materials are quite mobile, the bar form is not thought to be present: the form develops on the falling stage, and the bar is dissected on the lowest stage. The various bar forms have a related diagnostic structure which determined from the nature and distribution of the stratification types, and the manner of their deposition on the falling flow stage. The exposed (supra-platform) parts of lateral bars have a side and longitudinally filled inner channel which comprises fine sediment. Medial bars have converging cross-strata in sands which are overlain partly by gravels in an upward coarsening sequence.

Lateral migration of the channel zone, brought about by the preferential bar accretions to one side, results in building of a lithosome, the facies structure of which is partly determined by the nature of the bar accretions. As the channel migrates the abandoned bars may be covered with fine sediments so as to build up an upward fining sequence. The nature of the fine overbank part of the braided stream cycle is as complex as the coarse lower, so that a highly variable vertical sequence of lithological types is likely to be constructed.

Upward fining cycles are also built by some fine braided streams but they are the product of a single flood rather than lateral channel migration. Braided stream deposits are distinguishable from those of small, coarse meandering stream deposits by the presence of inner accretionary banks in the latter, and inner channels in the former.

Numerical taxonomic methods are applied to the Encrinurinae (Trilobita). Thirty-five cranidial and pygidial attributes are defined of which some (e.g. those relating to the glabellar tubercles) are specific to encrinurines and cybelines while others have application to non-encrinurids. Of nearly 200 named species of Encrinurinae only about one-third have been described in the literature in sufficient detail for their cranidial and pygidial attribute states be coded: a much smaller proportion of species is well enough known for librigenal, rostral, hypostomal and thoracic attributes to be coded. A comprehensive numerical taxonomy of the subfamily must therefore await detailed description of most of the known species. A list is given of encrinurine species in need of re-investigation. Meanwhile, 63 species can be coded for 34 attributes, and 75 species for 28 attributes. Principal components analysis is used to produce ordinations of these species. The minimal spanning tree superimposed on the ordination suggests possible evolutionary lineages. Ordination of the attributes is by non-linear mapping based on a distance matrix derived from the inter-attribute correlation matrix by the transformation (tanh−1 |r|)−1. The ordinations compare well with subjective assessments of relations and phylogenies.

Eurypterids of the Superfamily Stylonuroidea Diener 1924 sensu Størmer (1974, 373) from the Pentland Hills, Midlothian, are redescribed and the evidence which these forms may give concerning the life environment of the Gutterford Burn Eurypterid Bed (Upper Llandovery), from which most of them have been obtained, is considered. Five species are recognised. Parastylonurus ornatus (Laurie) is redescribed with special reference to the organs of locomotion and reproduction. A new form from the Gutterford Burn is described as Parastylonurus hendersoni sp. nov. Stylonurus macrophthalmus Laurie is designated the type species of the new genus Hardieopterus and Stylonurus knoxae Lamont as the type species of the new genus Lamontopterus. The unique holotype of Lamontopterus knoxae is of morphological interest in showing evidence of the gut.

Stylonuroids from other Scottish Silurian localities are also considered. Two new species, Brachyopterella ritchiei sp. nov. from Seggholm and Hardieopterus (?) lanarki sp. nov. from the Logan Water are described. Stylonurella spinipes (Page) is redescribed and this species is reported for the first time from Seggholm.

The new information provided by this study has raised a number of problems of classification and the criteria upon which eurypterids are classified, and particularly those applied to the stylonuroid eurypterids, are re-assessed and a new classification of the Stylonuroidea proposed. This has required the description of new taxa; a new family, the Parastylonuridae, is proposed to accommodate Parastylonurus, Hardieopterus and probably Lamontopterus which share a unique combination of characters of the prosomal appendages and metastoma. A new genus, Kiaeropterus, is described to accommodate certain species previously assigned to Stylonurella but displaced from that genus as now emended.

A reconstruction of Parastylonurus ornatus in the walking position is attempted and has resulted in the recognition of a new generation of problems related to the functional morphology of these animals. Criteria which may appropriately be used in deciding the manner in which they walked and the posture which they adopted when doing so, are discussed. It is suggested that, in response to certain anatomical and physiological constraints which otherwise would have rendered the animals unstable in the walking position, the post-abdomen and telson have been specialised as hydrodynamic structures for monitoring water movements and vectoring the animal for maximum walking efficiency. The significance of this hypothesis is discussed in relation to the functional morphology and evolution of other members of the Stylonuroidea.