1. An account has been given of the composition and weights of nutrients in 3200 wheat plants from seven weeks before ear emergence until the crop was harvested. The distribution of the plant nutrients, and the rate of assimilation and translocation of these nutrients, have been discussed.

2. In the case of the whole plant, the percentage of nutrients in the dry matter decreased from the time of the first sampling, with the exception of silica, which had a tendency to remain constant throughout. In the case of the ear, diminution in the percentage of nutrients in the dry matter was observed from a fortnight after ear emergence until harvest.

3. Marked increases in the weights of all nutrients in the whole plant were noted until ear emergence, from then onwards assimilation was much slower. The plant attained its maximum quantities of the nutrients in the following order: potash seven weeks, lime and chlorine five weeks, nitrogen three weeks, carbon, phosphoric acid and silica two weeks, before harvest.

Assimilation of these had, therefore, ceased at the above stated times, although transference to the ear proceeded until within about a week before harvest.