A new crustacean, Castracollis wilsonae is described from a loose block of the Early Devonian Rhynie chert, found in the vicinity of Rhynie, Aberdeenshire, Scotland. It differs markedly from Lepidocaris rhyniensis, Scourfield 1926, the only other crustacean found in the Rhynie chert.

The material comprises complete individuals up to 8 mm long and fragmentary remains. The head is normally damaged and detached from the body, and damage to the anterior of the trunk suggests the specimens are exuviae. The head appears domed with a labrum, robust mandibles, and long biramous antennae. A few specimens exhibit a probable detached cephalo-thoracicshield. The trunk is multi-segmented, comprising similar ring-like somites. The thorax variably comprises up to 26 segments. The anterior 11 segments possess similar long, phyllopodous appendages, the remainder variably possess 10 to 15 phyllopodous appendages with a disposition of one per somite. The abdomen comprises up to 28 apodous segments. The posterior of the body comprises a telson with two furcal rami.

The crustacean is most probably a calmanostracan branchiopod. It occurs associated with Lepidocaris, charophytes, cyanobacteria and coprolites within a ‘clotted’ chert texture, indicating subaqueous deposition, most probably in a temporary freshwater pool in an area of surficial hydrothermal activity.

Metamorphosed abyssal ocean-floor tholeiitiic rocks, little affected by crustal contamination and probably formed at a spreading ridge, occur as a dyke complex in banded gabbros with oceanic affinities and as small tectonic lenses in serpentinised peridotite. Igneous textures and mineral assemblages have been partly or completely replaced in both dykes and gabbros. Metamorphic development of clinopyroxene and very Ti-rich hornblende at 900-1000°C was followed by a low temperature actinolite-chlorite assemblage and then by pectolite-bearing veins. Geochemical variations resulting from magmatic and metasomatic processes have been distinguished and plots of major and trace elements used to establish tectonic environment. The very high temperature metamorphic mineral growth associated with beerbachite formation indicates that the dyke rocks were in a high heat-flow environment for a considerable time, presumably in the vicinity of a spreading ridge. Obduction of a very hot slab of oceanic lithosphere accounts for the superimposed mineral assemblages.

Spatially associated pillow lavas have been affected by crustal contamination and are not comagmatic with the dykes. There are petrochemical characters indicating that some of the lavas are ocean-floor basalts. For other lavas, features suggesting emplacement in a continental or an island-arc environment could also be consistent with development in a marginal basin. However, there is so much variation in composition of pillows from margin to centre, from small to large and from metasomatic activity, that their compositional fields cannot be used for discrimination of basaltic type with the same confidence as for the dykes. Accordingly, it is suggested that the petrochemistry of sheeted dyke complexes, rather than of pillow lavas, be used for this purpose in the study of ophiolites.

I am indebted to the kindness of Professor Ramsay, Director-General, and of Professor Geikie, Director of the Scottish Branch of the Geological Survey of Great Britain, for the privilege of examining and describing a remarkable collection of fossil fish-remains from the Lower Carboniferous rocks (Calciferous Sandstone Series) of Eskdale and Liddesdale. Most of the specimens were collected by Mr Arthur Macconochie, one of the collectors attached to the Scottish Geological Survey; and Mr Walter Park of Brooklyn Cottage, Langholm, has also willingly co-operated in the search, so far as the district of Eskdale is concerned. I have myself also had the pleasure of twice visiting Eskdale, along with Mr Macconochie and Mr B. N. Peach, and on these occasions I obtained a few specimens for my own collection.

A Histological investigation has been made of growth and repair of periostracum in Mytilus edulis. Physical rather than chemical differences in the processes of secretion can account for the variations in the structure of periostracum which appear during normal growth and in that which follows damage. Amœbocytes take part in the production of periostracum by supplying material for the secretory cells. Reasons for the presence of other granular cells near the site of secretion are also suggested. The periostracum is secreted solely by the epithelium of the inner surface of the outer mantle fold, although initially it adheres to the epithelium of a process of the middle fold. Consideration of the nature of the connection of the periostracum with its retractor muscles leads to the hypothesis that the cells of both epithelia move with the periostracum towards the mantle edge. The tissue where the two epithelia meet appears to be meristematic.

The results are presented of a detailed petrological reconnaissance of a group of picritic minor intrusions in the Hebrides. A substantial amount of new factual data is subjected to a unified treatment as a basis for reference and discussion. Olivine phenocrysts are not appreciably zoned and there is no evidence that they have a reaction relation with the liquid represented by the groundmass. Variations in the size and amount of olivine in individual intrusions are examined in detail and attributed to composite intrusion of differentiated material. A distinctive non-porphyritic facies found in several sills and in one dyke is chemically analyzed. Four analyses from widely separated localities establish this facies as a remarkably invariant, eucritic rock-type. The composition of the groundmass of the picritic rocks is variable and there is no evidence whatever of the participation of basaltic magma in their formation. Although no attempt is made to explain the new data in detail, a comprehensive working hypothesis is formulated. The origin of such picritic intrusions is believed to be due to selective fusion of pre-existing ultrabasic rock. Liquid more basic than normal basalt magmas can be formed by this process. Some re-precipitation of olivine may have preceded final emplacement of a magnesia-rich liquid which contained xenocrysts, mainly of olivine, from the source rock.

The Devonian actinopterygian Mimia Gardiner & Bartram (1977) of the Upper Devonian (Frasnian) Gogo Formation of Western Australia, one of the most completely known of all Palaeozoic ray-finned fishes, is renamed Mimipiscis nom. nov. due to preoccupation of the former genus by the butterfly Mimia Evans (1953). Recently acquired data, including the description of newly prepared fossil material, has revealed the presence of a second species in this formerly monotypic genus, as well as previously unreported features of the tail, parasphenoid and ontogenetic variability of type species, Mimipiscis toombsi (Gardiner & Bartram 1977). The second form, Mimipiscis bartrami sp. nov., differs from the contemporary M. toombsi in details of the body shape, squamation, snout, suboperculum and parasphenoid. Phylogenetic analyses recover Mimipiscis and Gogosardina as sister genera within a monophyletic Mimiidae, a clade restricted to the Late Devonian of Western Australia.

The material for this work was that collected and prepared by Professor J. Graham Kerr in the Chaco (Lepidosiren), and by Budgett from the Gambia (Protopterus). The process of development follows nearly the same course in the two genera, and where differences occur they are mainly such as are to be expected from a comparison of the adult skulls. While the series of stages of Lepidosiren left no unfilled gaps, the material was scantier in the case of Protopterus. Except where otherwise stated, the detailed descriptions were worked out in Lepidosiren, but whenever I was unable to verify that the process was substantially the same in the other genus, the fact has been mentioned. The stages are numbered in correspondence with the stages figured by Graham Kerr (Phil. Trans. Roy. Soc, B., vol. cxcii.). The Protopterus stages are numbered the same as the corresponding ones in the other genus. As far as possible, the nomenclature adopted by Gaupp* has been adhered to.

All trilobites known from the lower and upper Balclatchie and lower Ardwell groups of the Girvan district are listed and frequencies recorded. Eight new species are described, a number of generic assignments are new, and hitherto undescribed parts are illustrated. Of the 60 named species, 19 occur in the lower and upper Balclatchie and lower Ardwell groups; the upper Balclatchie Group yields the largest number of species. The assemblage is generally indicative of outer shelf conditions of deposition; a deeper water association at certain Balclatchie Group localities consists of 5 species identical with, or closely related to, species from the Liberty Hall facies of the lower Edinburg Formation of Virginia, U.S.A. This deeper water element constitutes a significantly larger proportion of the trilobite assemblages at certain localities than it does at others. A further 8 species from the Balclatchie Group are allied to lower Edinburg Formation forms, thus confirming the evidence of the brachiopods regarding the strong North American affinities of the fauna. The Balclatchie conglomerate is considered to be the result of a catastrophic event accompanied by slumping near the shelf edge causing the instantaneous entombment of the fauna.

Pentastomida, tongue worms, are a taxon of about 130 species of parasites, living exclusively in the respiratory tracts of vertebrates. Three-dimensionally preserved Upper Cambrian larvae already demonstrate a high degree of adaptation to parasitism, striking morphological conservatism, and a high diversification by the Late Cambrian, thereby suggesting a likewise diversified host group. Not least due to their highly modified morphology, the systematic affinities of pentastomids remain controversial. The two major alternatives place the group as either close to branchiuran crustaceans or as stem-lineage derivatives of the Euarthropoda. To this set of Cambrian fossil representatives of the pentastomids we can add a new form from Lower Ordovician boundary beds from Sweden, most likely reworked from Upper Cambrian horizons. Based on this new species, named Aengapentastomum andresi gen. et sp. nov., and the available information about fossil and Recent pentastomids, we review the diverging ideas on the systematic position of this fully parasitic taxon.

Leucogranites are typical products of collisional orogenies. They are found in orogenic terranes of different ages, including the Proterozoic Trans-Hudson orogen, as exemplified in the Black Hills, South Dakota, and the Appalachian orogen in Maine, both in the USA, and the ongoing Himalayan orogen. Characteristics of these collisional leucogranites show that they were derived from predominantly pelitic sources at the veining stages of deformation and metamorphism in upper plates of thickened crusts. Once generated, the leucogranite magmas ascended as dykes and were emplaced within shallower parts of their source sequences. In these orogenic belts, there was a strong connection between deformation, metamorphism and granite generation. However, the heat sources needed for partial melting of the source rocks remain controversial. Lack of evidence for significant intrusion of mafic magmas necessary to cause melting of upper plate source rocks suggests that leucogranite generation in collisional orogens is mainly a crustal process.

The present authors evaluate five types of thermal models which have previously been proposed for generating leucogranites in collisional orogens. The first, a thickened crust with exponentially decaying distribution of heat-producing radioactive isotopes with depth, has been shown to be insufficient for heating the upper crust to melting conditions. Four other models capable of raising the crustal temperatures sufficiently to initiate partial melting of metapelites in thickened crust include: (1) thick sequences of sedimentary rocks with high amounts of internal radioactive heat production; (2) decompression melting; (3) thinning of mantle lithosphere; and (4) shear-heating. The authors show that, for reasonable boundary conditions, shear-heating along crustal-scale shear zones is the most viable process to induce melting in upper plates of collisional orogens where pelitic source lithologies are usually located. The shear-heating model directly links partial melting to the deformation and metamorphism that typically precede leucogranite genera

Fluid dynamic modelling of crystallising calc-alkalic magma bodies has predicted that differentiated liquids will ascend as boundary layers and that accumulation of these buoyant liquids near chamber roofs will result in compositionally stratified magma chambers. This paper reports physical features in La Gloria Pluton that can be interpreted as trapped ascending differentiated liquids. Leucogranitic layers decimetres thick, which are locally stratified, are trapped beneath overhanging wall contacts. The same felsic magmas were also preserved where they were injected into the wall rocks as dykes and as large sill complexes. These rocks do not represent differentiated magmas produced by crystallisation along the exposed walls because the felsic layers occur at the wall rock contact, not inboard of it. Rather, we speculate that evolved felsic liquids are generated by crystallisation all across the deep levels of chambers and that initial melt segregation occurs by flowage of melt into tension fractures. Melt bodies so formed may be large enough to have significant ascent velocities as diapirs and/or dykes. The other way in which the leucogranite occurrence is at variance with the convective fractionation model is that the ascending liquids did not feed a highly differentiated cap to the chamber, as the composition at the roof, although the most felsic in this vertically and concentrically zoned pluton, is considerably more mafic than the trapped leucogranitic liquids. This suggests that these evolved liquids were usually mixed back into the main body of the chamber. Backmixing may be general in continental-margin calc-alkalic magmatic systems, which, in contrast to those in intracontinental settings, rarely produce volcanic rocks more silicic than rhyodacite. That the highly differentiated liquids are preserved at all at La Gloria is a result of the unusual stepped nature of the contact and the entirely passive mode of emplacement of the pluton, which, in contrast to ballooning in place, does not result in wall zones being “scoured”.

Examination of living embryos has shown that the heart is a functionally active organ from a very early stage of its development. At all periods of life the result of the functional activity is in essentials the same, viz. the propulsion of the blood in a definite direction through the heart into the vessels arising from it; but the mechanism for effecting this propulsion undergoes profound alterations, and the heart becomes transformed from a simple continuous tube, destitute of valves, whose walls contract in a rhythmic peristaltic wave, into a complex four-chambered organ, divided into right and left portions, which are ultimately completely separated from one another, possessing valves, and contracting not in a peristaltic wave but in alternating consecutive contractions of the atria and ventricles of the right and left sides simultaneously. Coincidently with the changes in the heart itself, profound alterations occur in the vessels leading to and from the heart. In this combination of simultaneous development and functional activity the heart differs from the other organs of the body, and hence its development presents special problems involving the function as well as the structure of the different parts. Our knowledge of the development of the heart in man cannot yet be said to be complete.

The dermal skull roof of Acanthostega gunnari Jarvik is described in detail, giving new information on the anatomy, and new reconstructions of the skull in dorsal, lateral and occipital views, as well as sections through the skull at two points. The types of suture are compared with those of temnospondyls and discosauriscids, comparable animals in terms of habit and skull shape. The skull-bones of Acanthostega are much more closely integrated with each other than those of the two later groups. However, regions of relatively less firmly and more firmly sutured regions are comparable in position. This may be connected with having the lateral lines accommodated in tubes in Acanthostega rather than grooves, or some other property of the dermal bone in the later groups that reduced the need for stronger integration. The post-orbital region of the skull (skull table) is the most strongly integrated region, possibly connected with the need for attachment of occipital musculature. The skull table and dentition are the most labile of cranial features among Devonian tetrapods, whereas the snout, cheek, palate and infradentary regions remain conservative. These are for the most part the lateral-line-bearing regions of the skull. The specimens of Acanthostega found at one site on Stensiö Bjerg may form part of a year-group that had perhaps come together for spawning.

Orthopyroxene- and kyanite-lineage eclogites are distinguished on the basis of silicate and carbonate petrography and chemistry. Different textural and chemical types of magnesite, dolomite and calcite can be satisfactorily attributed to one of the three sequential metamorphic stages of eclogite history at anhydrous eclogite facies, hydrous eclogite facies and amphibolite facies. The chronological sequence of development of magnesite, dolomite and calcite matches the carbonate sequence found experimentally in synthetic peridotite-CO2-H2O systems during decompression. Magnesite and dolomite show reasonably regular Fe/Mg distribution relationships with silicates.

A new geothermobarometer based upon is provisionally calibrated. It yields T estimates for given P very similar to those given by the geothermobarometer, e.g. 759–829°C at 20 kb and 803–894°C at 40 kb. P is tentatively estimated at 30±10, 23±6 and 10 ± 3 kb respectively for the three metamorphic stages.

Fluid conditions after equilibration during each stage were either low or fluid-absent, but not low . A unique quartz-magnesite-dolomite vein represents an influx of CO2 in the P-T regime of instability of CO2 + pyroxene. Tectonic emplacement by deep-level obduction of diverse foreign source materials remains the petrogenetic model which best fits the data for these eclogites.

The main purpose of the following investigation is to improve the existing Approximate Theory of Beams by bringing it into closer relationship than has hitherto been done with exact solutions of the fundamental differential equations of equilibrium. In its aim, method, and results the paper closely resembles one already published in these Transactions on the corresponding Theory for Plates (vol. xli., 1904). It is less comprehensive than the latter paper, as it deals only with a beam of circular section; but the analysis is of such a nature that the chief results can be extended almost immediately to a beam of any form of section, on the assumption merely of certain Existence Theorems.

Conchyliocarida, from the Jurassic of France, have a conch-like carapace enclosing the whole body. The number of their somites, identified by the visible appendages, excludes them from Malacostraca. Caridean-like characters, including the large lamellar gills, bring them nearer to Malacostraca, end exclude them from other Classes. Their large functional compound eyes and the morphology of their limbs suggest a benthic life and predatory behaviour.

Conchyliocarida are represented by several forms at La Voulte sur Rhône, which are compared and contrasted to related forms elsewhere.

The currently accepted cladogram of spider phylogeny and palaeontological data are used to evaluate spider family richness through geological time. A significantly more diverse spider fossil record is predicted than observed. The predicted rate of spider family diversification is considered more accurate because of its close similarity at 0 Ma to the number of extant families. Predicted spider family palaeodiversity is compared with insect family palaeodiversity to investigate whether spiders track insects through geological time. At the family level, the insects, and observed and predicted spider fossil records show an exponential increase over time, the pattern typical of a radiating group. No significant differences are observed in the rates of change in the slopes, and hence rate of diversification of spiders and insects over time. This suggests that spiders probably co-radiated alongside the insects, with the major radiations of both groups occurring at least 100 Ma before the origin of angiosperms.

A new reconstruction of the skull of Crassigyrinus scoticus (Lydekker) is provided, based on newly prepared material. A revised interpretation of the naris shows it in most respects to be like that of other Carboniferous tetrapods, lacking both an anterior tectal and a lateral rostral. However it does incorporate an enlarged septomaxilla, and part of the dorsal surface of the vomer is also visible within the naris of most specimens. Prominent paired buttresses run along the midline of the snout, and a single bowl-shaped interpremaxillary fenestra pierces the snout and palate but is not confluent with the naris or choana. A preopercular is probably absent. A new cladistic analysis places Crassigyrinus with the Viséan genus Whatcheeria, and puts those two taxa as a sister-clade to the anthracosaurs Proterogyrinus and Pholiderpeton. Temnospondyls form a clade with loxommatids, and these two arms form a dichotomy crownward to Greererpeton, Ichthyostega and Acanthostega which are successive stemward plesions.

Mafic rocks in the Ryoke belt, the Cretaceous granitic province in Southwest Japan, occur in two modes: (1) as mafic dykes and pillow-shaped enclaves, and (2) as isolated kilometresized bodies of gabbroic cumulate. The dykes and pillows have fine-grained textures with thin radiating plagioclase laths, indicative of quenching. The gabbroic cumulates are predominantly coarse-grained and commonly lithologically layered.

SHRIMP zircon U-Pb ages of both types of mafic rocks are in the range 71–86 Ma, late Cretaceous. The mafic rocks become younger eastwards, matching the along-arc age trend of the associated Cretaceous granites (Nakajima et al. 1990). Both types of mafic rocks were apparently generated during the same magmatic event that produced the Ryoke/San-yo granites. The mafic dykes and pillows are aphyric basaltic-andesites to andesites (SiO2 54–60 wt.%), with microphenocrysts of biotite and hornblende. They have a composition which is similar to mafic rocks from the northern Sierra Nevada, and also to medium-K calc-alkaline rocks from present-day arc volcanics. The gabbroic cumulates are mostly pyroxene-hornblende gabbros (SiO2 43–52 wt.%). Their bulk-rock chemical compositions are mostly unlike any magma compositions.

Both types of mafic rocks from the Ryoke belt have relatively high 87Sr/86Sr initial ratios (SrI), 0·7071–0·7097, which are similar to those of the associated granites. The granites were formed either by fractional crystallisation of the mafic magmas, or by partial melting of newly formed mafic rocks at depth. The high SrI indicates that the mafic magmas were derived from enriched mantle or mixed with enriched crustal materials. Even if the mixing occurred between primitive basaltic magma and metasedimentary rocks, then the basaltic andesite–andesite magmas must have contained more than 60% mantle-derived components. The Cretaceous magmatism in Southwest Japan represents a major episode of crustal growth by additions from the upper mantle in an arc setting.

Forty-one species assigned to thirty-seven named genera of trilobites are described from the Tangtou Formation at Tangshan and Lunshan, in Jiangsu Province, SE China. Nankinolithus nankinensis Lu, the Zone fossil of the upper part of the Formation, is abundant. Many Tangtou Formation species occur also in the Huangnehkan, Linhsiang and Chiencaokou formations, and all are considered to be of early Ashgill age. The trilobites are closely related to those of the Staurocephalus clavifrons Zone (Rawtheyan) of Poland, demonstrating the long duration and wide extent of deeper water faunas. There are also close links with the Ashgill faunas of Central Asia.

One new species, Amphitryon cheni, is established. The following morphological features are described: a small remopleuridine hypostome is unlike any other in construction; Nileus transversus retains genal spines to full grown size in some specimens, but loses them in others; the hypostome lacks the posteromedian cusp of other species. The hypostome of Paraphillipsinella is strongly styginid in conformation, and unlike that of Phillipsinella. Telephina convexa is the only species known with three spines on the librigena. The arrangement of pits in Nankinolithus nankinensis, the type species, is described, including illustrations of the irregularities which occur in every fringe. A small bulb at the tip of the hypostome of Ovalocephalus (= Hammatocnemis) bears two backwardly pointing spinules, one above the other, on the vertical posterior margin. Three types of protaspides, and numerous meraspid parts are described.